In my opinion, you've got two things wrong , firstly, see this Default Receive Connectors:
Default Fronted Server does not work for internal mail flow. Also, the description you mentioned might be too old, we don't have hub server since Exchange 2013. In case it is modified to "This service proxies SMTP connections inbound/outbound to mailbox server", it's still not talking about the internal mail flow things, tell me if I misunderstand your meanings.
Secondly, see this transport pipeline:
When mail routing between exchange servers, front end transport service is not involved. The key connector for internal mail flow is named "Default <servername>" and the port is 2525, for further information see Default Receive connectors in the Transport service on Mailbox servers.
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