A family of Microsoft relational database management systems designed for ease of use.
First off, most of us really dislike the "Lookup Field" datatype. It sounds like you're wanting to use a Combo Box in a Table - this is very limited and often confusing. See The Access Web for a critique. Your Country field in mutationtable should NOT be a Lookup Field; instead it should be a Number, Long Integer field matching the ID of the Countries table. In a Form you can thus store the ID (keeping the computer happy) while displaying the country name (keeping the user happy).
A Combo Box will autocomplete if its autocomplete property is turned on, and the field (Country name in this case) is the first visible column. It works well if you have the combo box's Row Source query sorted by the name field (so it will find Algeria before Azerbaijan for example).