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Can guest users in my organization change their usernames and email addresses in the same way as members?
No, as their properties are managed in their "home" tenant, but the sync process is one-time only, upon creation. If changes need to be reflected on the "resource" side, someone with admin access on the resource tenant will have to do it.
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Hi @Anonymous
In Office 365, the guest users are represented by a mail user objects, so if you want to change email address for the existing guest user account, you can use the Exchange powershell cmdlet Set-MailUser.
Set-MailUser <mail_alias_of_guest_user> -WindowsEmailAddress <new email addresss>