A family of Microsoft relational database management systems designed for ease of use.
First, It is not recommended that you do lookups on the table level. This can cause more problems then its worth. All lookups should be done using list controls on a form.
Second, what you are referring to can only be done on a form. You would set the RowSource of your combo to look like:
SELECT CompanyID, CompanyName
FROM tblCompany
WHERE CompanyName LIKE Forms!formname!textboxname & "*;"
So whatever letter you type in the textbox, will filter the list of companys.